Marine biology and ecology, sclerochronology, scientific diver

Associate Professor, Habilitation
Université de Bretagne Occidentale


LEMAR Laboratory




+33 (0) 2 98 49 86 47



  • 2019 – Habilitation thesis, University of Western Brittany
  • 2005 – Ph.D., Biological Oceanography, University of Western Brittany
  • 2001 – Master of Science in Marine Ecology, University of Western Brittany
  • 1998 – Bachelor’s Degree in Biology, University of Western Brittany & University of Rennes 1

I am working as Associate Professor at the University of Western Brittany since 2009, after a Ph.D. thesis carried out in New Caledonia, and two post-doctoral stays in the USA with Jim Cloern (US Geological Survey, Menlo Park, CA) then in Germany with Bernd Schöne (Johannes Gutenberg-Universität, Mainz). My research activities mainly focus on the description and understanding of the ecological functioning of coastal ecosystems.

Marine ecologist by training, I began to combine these skills with the thematic field of biogenic carbonate geochemistry during my Ph.D., and then immersed myself in the analysis of the composition of aquatic mollusc shells using state-of-the-art techniques generally used in geosciences (laser ablation, mass spectrometry ICP-MS and IR-MS). The coupling between these geochemical methods and the autoecology of mollusks allowed me to calibrate various proxies of environmental parameters archived in shells (e.g. temperature, phytoplankton dynamics), then to develop paleo-ecological and paleo-environmental reconstructions over periods ranging from a year (with a daily temporal resolution) to several centuries (annual resolution). Today, the core of my research activities still deals with extracting, analyzing and understanding structural (variation of growth increment width) and geochemical (variation of stable isotope ratios and elemental concentrations) proxies archived in mollusc shells from coastal ecosystems (Europe, Arctic, Antarctic, Pacific, West Africa). This scientific field is called “sclerochronology”, the aquatic equivalent of dendrochronology (study of tree rings). By extension, sclerochemistry is the sub-discipline of sclerochronology dealing with microchemical analyses performed in these structures.

The main objectives of my research activities are (i) to get information on life-history traits (of aquatic molluscs (bivalves and gastropodes), (ii) to assess past and present variability of environmental conditions (temperature, salinity, primary production, pollutions, etc.), and ultimately (iii) to describe and understand the ecological functioning of temperate, tropical and polar coastal ecosystems.

I am also a scientific diver, holding the Hyperbaric Aptitude Certificate (class 1, mention B), the Scientific Dive Leader qualification, and the Air Diluent Closed Circuit Rebreather (rEvo) certification.

Main scientific collaborations:

  • Prof. Melita Peharda, Institute of Oceanography and Fisheries, Split, Croatia
  • Prof Bernd R. Schöne, Johannes Gutenberg-Universität, Mainz, Germany

Annual growth lines in Arctica shells

Besides my research activities, I also teach at the University of Western Brittany, both at the Faculty of Sciences & Technology (Bachelor of Biology) and at the European Institute for Marine Studies (IUEM – Master Marine and Coastal Sciences). I teach (or used to) zoology, general ecology, functional biology, population dynamics, methods and practices in ecology, as well as a pinch of taxonomy to undergraduate students. At the Master 1 level, I teach population biology and more particularly its evolutionary aspects, conservation biology and ecology of polar and tropical environments. Finally, I initiate Master 2 students to environmental biological records, a teaching unit consisting of an introduction to sclerochronology and sclerochemistry of aquatic organisms. I also act as a tutor for graduate student internships, and supervise the research work of master and PhD students.

Ph.D. students

  • Mr Valentin Siebert, 2019-2023: Bivalve shell-based, high-resolution multi-proxy reconstruction of marine primary production – Pecten maximus, Bay of Brest
  • Ms Justine Doré, 2017-2020: Sclerochronology and geochemistry of Northwestern Atlantic bivalves: Exploration of environmental proxies of global changes
  • Mr. Pierre Poitevin, 2015-2018: Sclerochronology and sclerochemistry of three soft bottom bivalve species from Saint-Pierre et Miquelon
  • Ms Amy Featherstone, 2014-2017: Construction of long annually-resolved shell-based chronologies using Glycymeris glycymeris (Bivalvia; Glycymeridae) from the Bay of Brest, France
  • Ms. Clémence Royer, 2011-2014: Palaeoecological reconstruction of the impact of anthropogenic activities on aquatic ecosystems of the Bay of Brest and its watersheds: sclerochronological and geochemical investigations on shells of the freshwater pearl mussel (Margaritifera margaritifera) and the dog cockle (Glycymeris glycymeris)

Undergraduate and graduate students

  • Ms Chloé Malirat, 2022, University of Bordeaux
  • Mr Samuel Martin, 2021, University of Lille 1
  • Mr Clément Gauchot, 2021, University of Western Brittany
  • Ms Marie Le Bagousse, 2021, University of Western Brittany
  • Mr Pierre-Antoine Renaud, 2020, University of Southern Brittany
  • Mr Louis Quichaud, 2020, ENS Lyon
  • Ms. Elena Bogdaniuk, 2018, University of Western Brittany
  • Mr. Valentin Siebert, 2018, University of Western Brittany
  • Mr Johan Vieira, 2017, University of Bordeaux
  • Ms Julie Bellec, 2016, University of Bordeaux
  • Mr Sylvain Menneteau, 2016, University of Western Brittany
  • Ms Sophie Sabarot, 2015, University of Pau
  • Ms Justine Doré, 2015, University of Aix-Marseille
  • Mr Quentin Morel, 2014, University of Western Brittany
  • Ms Véronique Pérez, 2012, Université du Québec à Rimouski
  • Mr Adhitya Kusuma Wardana, 2012, Diponegoro University (Semarang, Indonesia) & University of Western Brittany
  • Ms Clémentine Le Jouan, 2012, University of Western Brittany
  • Ms Blandine Gaillard, 2011, University of Western Brittany
  • Mr Romain Lavaud, 2010, University of Western Brittany
  • Ms Clémence Royer, 2010, University of Western Brittany

55. Carré M., Quichaud L., Camara A., Azzoug M., Cheddadi R., Ochoa D., Cardich J., Pérez A., Salas-Gismondi R., Thébault J. & Thomas Y. (2022). Climate change, migrations, and the peopling of sine-Saloum mangroves (Senegal) in the past 6000 years. Quaternary Science Reviews 293: 107688.
doi: 10.1016/j.quascirev.2022.107688

54. Fröhlich L., Siebert V., Huang Q., Thébault J., Jochum K.P. & Schöne B.R. (2022). Deciphering the potential of Ba/Ca, Mo/Ca and Li/Ca profiles in the bivalve shell Pecten maximus as proxies for the reconstruction of phytoplankton dynamics. Ecological Indicators 141: 109121.
doi: 10.1016/j.ecolind.2022.109121

53. Fröhlich L., Siebert V., Walliser E.O., Thébault J., Jochum K.P., Chauvaud L. & Schöne B.R. (2022). Ba/Ca profiles in shells of Pecten maximus – A proxy for specific primary producers rather than bulk phytoplankton. Chemical Geology 593: 120743.
doi: 10.1016/j.chemgeo.2022.120743

52. Thébault J., Jolivet A., Waeles M., Tabouret H., Sabarot S., Pécheyran C., Leynaert A., Jochum K.P., Schöne B.R., Fröhlich L., Siebert V., Amice E. & Chauvaud L. (2022). Scallop shells as geochemical archives of phytoplankton-related ecological processes in a temperate coastal ecosystem. Limnology and Oceanography 67(1): 187-202.
doi: 10.1002/lno.11985

51. Ricardo F., Mamede R., Bruzos A.L., Díaz S., Thébault J., Ferreira da Silva E., Patinha C. & Calado R. (2022). Assessing the elemental fingerprints of cockle shells (Cerastoderma edule) to confirm their geographic origin from regional to international spatial scales. Science of the Total Environment 814: 152304.
doi: 10.1016/j.scitotenv.2021.152304

50. Doré J., Thébault J., Roy V., Dewilde F. & Chaillou G. (2021). Insight into shell growth dynamics of Mactromeris polynyma in the St. Lawrence Estuary using sclerochemistry. Marine Chemistry 234: 103987.
doi: 10.1016/j.marchem.2021.103987

49. Siebert V., Poitevin P., Chauvaud L., Schöne B.R., Lazure P. & Thébault J. (2021). Using growth and geochemical composition of Clathromorphum compactum to track multiscale North Atlantic hydro-climate variability. Palaeogeography, Palaeoclimatology, Palaeoecology 562: 110097.
doi: 10.1016/j.palaeo.2020.110097

48. Trofimova T., Alexandroff S.J., Mette M., Tray E., Butler P.G., Campana S., Harper E., Johnson A.L.A., Morrongiello J.R., Peharda M., Schöne B.R., Andersson C., Andrus C.F.T., Black B.A., Burchell M., Carroll M.L., DeLong K.L., Gillanders B.M., Grønkjær P., Killam D., Prendergast A.L., Reynolds D.J., Scourse J.D., Shirai K., Thébault J., Trueman C. & de Winter N. (2020). Fundamental questions and applications of sclerochronology: Community-defined research priorities. Estuarine, Coastal and Shelf Science 245: 106977.
doi: 10.1016/j.ecss.2020.106977

47. Olivier F., Gaillard B., Thébault J., Meziane T., Tremblay R., Dumont D., Bélanger S., Gosselin M., Jolivet A., Chauvaud L., Martel A., Rysgaard S., Olivier A.-H., Pettré J., Mars J., Gerber S. & Archambault P. (2020). Shells of the bivalve Astarte moerchi give new evidence of a strong pelagic-benthic coupling shift occurring since the late 1970s in the NOW polynya. Philosophical Transactions of the Royal Society A 378: 20190353.
doi: 10.1098/rsta.2019.0353

46. Poitevin P., Chauvaud L., Pécheyran C., Lazure P., Jolivet A. & Thébault J. (2020). Does trace element composition of bivalve shells record ultra-high frequency environmental variations? Marine Environmental Research 158: 104943.
doi: 10.1016/j.marenvres.2020.104943

45. Doré J., Chaillou G., Poitevin P., Lazure P., Poirier A., Chauvaud L., Archambault P. & Thébault J. (2020). Assessment of Ba/Ca in Arctica islandica shells as a proxy for phytoplankton dynamics in the Northwestern Atlantic Ocean. Estuarine, Coastal and Shelf Science 237: 106628.
doi: 10.1016/j.ecss.2020.106628

44. Thibault M., Duprey N., Gillikin D.P., Thébault J., Douillet P., Chauvaud L., Amice E., Munaron J.-M. & Lorrain A. (2020). Bivalve δ15N isoscapes provide a baseline for urban nitrogen footprint at the edge of a World Heritage coral reef. Marine Pollution Bulletin 152: 110870.
doi: 10.1016/j.marpolbul.2019.110870

43. Featherstone A.M., Butler P.G., Schöne B.R., Peharda M. & Thébault J. (2020). A 45-year sub-annual reconstruction of seawater temperature in the Bay of Brest, France, using the shell oxygen isotope composition of the bivalve Glycymeris glycymeris. The Holocene 30(1): 3-12.
doi: 10.1177/0959683619865592

42. Poitevin P., Thébault J., Siebert V., Donnet S., Archambault P., Doré J., Chauvaud L. & Lazure P. (2019). Growth response of Arctica islandica to North Atlantic oceanographic conditions since 1850. Frontiers in Marine Science 6: 483.
doi: 10.3389/fmars.2019.00483

41. Peharda M., Thébault J., Markulin K., Schöne B.R., Janeković I. & Chauvaud L. (2019). Contrasting shell growth strategies in two Mediterranean bivalves revealed by oxygen-isotope ratio geochemistry: The case of Pecten jacobaeus and Glycymeris pilosa. Chemical Geology 526: 23-35.
doi: 10.1016/j.chemgeo.2017.09.029

40. Purroy A., Najdek M., Isla E., Župan I., Thébault J. & Peharda M. (2018). Bivalve trophic ecology in the Mediterranean: Spatio-temporal variations and feeding behavior. Marine Environmental Research 142: 234-249.
doi: 10.1016/j.marenvres.2018.10.011

39. Poitevin P., Thébault J., Schöne B.R., Jolivet A., Lazure P. & Chauvaud L. (2018). Ligament, hinge, and shell cross-sections of the Atlantic surfclam (Spisula solidissima): Promising marine environmental archives in NE North America. PLoS ONE 13(6): e0199212.
doi: 10.1371/journal.pone.0199212

38. Purroy A., Milano S., Schöne B.R., Thébault J. & Peharda M. (2018). Drivers of shell growth of the bivalve, Callista chione (L. 1758) – Combined environmental and biological factors. Marine Environmental Research 134: 138-149.
doi: 10.1016/j.marenvres.2018.01.011

37. Guédron S., Point D., Acha D., Bouchet S., Baya P.A., Tessier E., Monperrus M., Molina C.I., Groleau A., Chauvaud L., Thébault J., Amice E., Alanoca L., Duwig C., Uzu G., Lazarro X., Bertrand A., Bertrand S., Barbraud C., Delord K., Gibon F.M., Ibanez C., Flores M., Fernandez Saavedra P., Ezpinoza M.E., Heredia C., Rocha F., Zepita C. & Amouroux D. (2017). Mercury contamination level and speciation inventory in Lakes Titicaca & Uru-Uru (Bolivia): Current status and future trends. Environmental Pollution 231: 262-270.
doi: 10.1016/j.envpol.2017.08.009

36. Featherstone A.M., Butler P.G., Peharda M., Chauvaud L. & Thébault J. (2017). Influence of riverine input on the growth of Glycymeris glycymeris in the Bay of Brest, North-West France. PLoS ONE 12(12): e0189782.
doi: 10.1371/journal.pone.0189782

35. Bertucci A., Pierron F., Thébault J., Klopp C., Bellec J., Gonzalez P. & Baudrimont M. (2017). Transcriptomic responses of the endangered freshwater mussel Margaritifera margaritifera to trace metals contamination in the Dronne River. Environmental Science and Pollution Research 24: 27145-27159.
doi: 10.1007/s11356-017-0294-6

34. De Cesare S., Méziane T., Chauvaud L., Richard J., Sejr M., Thébault J., Winkler G. & Olivier F. (2017). Dietary plasticity in the bivalve Astarte moerchi revealed by a multimarker study in two Arctic fjords. Marine Ecology Progress Series 567: 157-172.
doi: 10.3354/meps12035

33. Purroy A., Šegvić-Bubić T., Holmes A., Bušelić I., Thébault J., Featherstone A. & Peharda M. (2016). Combined use of morphological and molecular tools to resolve species mis-identifications in the Bivalvia The case of Glycymeris glycymeris and G. pilosa. PLoS ONE 11(9): e0162059.
doi: 10.1371/journal.pone.0162059

32. McFarland K., Jean F., Thébault J. & Volety A.K. (2016). Potential impacts of blooms of the toxic dinoflagellate Karenia brevis on the growth, survival and juvenile recruitment of the non-native green mussel Perna viridis in southeastern United States. Toxicon 109: 94-102.
doi: 10.1016/j.toxicon.2015.11.017

31. Jolivet A., Chauvaud L., Thébault J., Robson A.A., Dumas P., Amos G. & Lorrain A. (2015). Circadian behaviour of Tectus (Trochus) niloticus in the southwest Pacific inferred from accelerometry. Movement Ecology 26:3.
doi: 10.1186/s40462-015-0054-5

30. Marchais V., Richard J., Jolivet A., Flye-Sainte-Marie J., Thébault J., Jean F., Richard P., Paulet Y.-M., Clavier J. & Chauvaud L. (2015). Coupling experimental and field-based approaches to decipher carbon sources in the shell of the great scallop, Pecten maximus (L.). Geochimica et Cosmochimica Acta 168: 58-69.
doi: 10.1016/j.gca.2015.07.010

29. Zeppilli D., Sarrazin J., Leduc D., Martinez Arbizu P., Fontaneto D., Fontanier C., Gooday A.J., Kristensen R.M., Ivanenko V.N., Sørensen M.V., Vanreusel A., Thébault J., Mea M., Allio N., Andro T., Arvigo A., Castrec J., Danielo M., Foulon V., Fumeron R., Hermabessiere L., Hulot V., James T., Langonne-Augen R., Le Bot T., Long M., Mahabror D., Morel Q., Pantalos M., Pouplard E., Raimondeau L., Rio-Cabello A., Seite S., Traisnel G., Urvoy K., Van Der Stegen T., Weyand M. & Fernandes D. (2015). Is the meiofauna a good indicator for climate change and anthropogenic impacts? Marine Biodiversity 45: 505-535.
doi: 10.1007/s12526-015-0359-z

28. Peharda M., Puljas S., Chauvaud L., Schöne B.R., Ezgeta-Balić, D. & Thébault J. (2015). Growth and longevity of Lithophaga lithophaga: what can we learn from shell structure and stable isotope composition? Marine Biology 162: 1531-1540.
doi: 10.1007/s00227-015-2690-0

27. Jolivet A., Asplin L., Strand Ø., Thébault J. & Chauvaud L. (2015). Coastal upwelling in Norway recorded in Great Scallop shells. Limnology and Oceanography 60: 1265-1275.
doi: 10.1002/lno.10093

26. Peharda M., Calcinai B., Puljas S., Golubić S., Arapov J. & Thébault J. (2015). Endoliths in Lithophaga lithophaga shells – Variation in intensity of infestation and species occurrence. Marine Environmental Research 108: 91-99.
doi: 10.1016/j.marenvres.2015.05.002

25. Jolivet A., Chauvaud L., Huchette S., Legoff C., Thébault J., Nasreddine K., Schöne B.R. & Clavier J. (2015). The ormer (Haliotis tuberculata): A new, promising paleoclimatic tool. Palaeogeography, Palaeoclimatology, Palaeoecology 427: 32-40.
doi: 10.1016/j.palaeo.2015.03.032

24. Biscéré T., Rodolfo-Metalpa R., Lorrain A., Chauvaud L., Thébault J., Clavier J. & Houlbrèque F. (2015). Responses of two scleractinian corals to cobalt pollution and ocean acidification. PLoS ONE 10(4): e0122898.
doi: 10.1371/journal.pone.0122898

23. Poulain C., Gillikin D.P., Thébault J., Munaron J.-M., Bohn M., Robert R., Paulet Y.-M. & Lorrain A. (2015). An evaluation of Mg/Ca, Sr/Ca, and Ba/Ca ratios as environmental proxies in aragonite bivalve shells. Chemical Geology 396: 42-50.
doi: 10.1016/j.chemgeo.2014.12.019

22. Lorrain A., Clavier J., Thébault J., Tremblay-Boyer L., Houlbrèque F., Amice E., Le Goff M. & Chauvaud L. (2015). Variability in diel and seasonal in situ metabolism of the tropical gastropod Tectus niloticus. Aquatic Biology 23: 167-182.
doi: 10.3354/ab00618

21. Artigaud S., Lavaud R., Thébault J., Jean F., Strand Ø., Strohmeier T., Milan M. & Pichereau V. (2014). Proteomic-based comparison between populations of the Great Scallop, Pecten maximus. Journal of Proteomics 105: 164-173.
doi: 10.1016/j.jprot.2014.03.026

20. Waeles M., Dulaquais G., Jolivet A., Thébault J. & Riso R.D. (2013). Systematic non-conservative behavior of molybdenum in a macrotidal estuarine system (Aulne-Bay of Brest, France). Estuarine, Coastal and Shelf Science 131: 310-318.
doi: 10.1016/j.ecss.2013.06.018

19. Pérez V., Olivier F., Tremblay R., Neumeier U., Thébault J., Chauvaud L. & Méziane T. (2013). Trophic resources of the bivalve, Venus verrucosa, in the Chausey archipelago (Normandy, France) determined by stable isotopes and fatty acids. Aquatic Living Resources 26: 229-239.
doi: 10.1051/alr/2013058

18. Thébault J. & Chauvaud L. (2013). Li/Ca enrichments in great scallop shells (Pecten maximus) and their relationship with phytoplankton blooms. Palaeogeography, Palaeoclimatology, Palaeoecology 373: 108-122.
doi: 10.1016/j.palaeo.2011.12.014

17. Royer C., Thébault J., Chauvaud L. & Olivier F. (2013). Structural analysis and paleoenvironmental potential of dog cockle shells (Glycymeris glycymeris) in Brittany, northwest France. Palaeogeography, Palaeoclimatology, Palaeoecology 373: 123-132.
doi: 10.1016/j.palaeo.2012.01.033

16. Lavaud R., Thébault J., Lorrain A., van der Geest M. & Chauvaud L. (2013). Senilia senilis (Linnaeus, 1758), a biogenic archive of environmental conditions on the Banc d “Arguin (Mauritania). Journal of Sea Research 76: 61-72.
doi: 10.1016/j.seares.2012.11.003

15. Chauvaud L., Patry Y., Jolivet A., Cam E., Le Goff C., Strand Ø., Charrier G., Thébault J., Lazure P., Gotthard K. & Clavier J. (2012). Variation in size and growth of the great scallop Pecten maximus along a latitudinal gradient. PLoS ONE 7(5): e37717.
doi: 10.1371/journal.pone.0037717

14. Tabouret H., Pomerleau S., Jolivet A., Pécheyran C., Riso R., Thébault J., Chauvaud L. & Amouroux D. (2012). Specific pathways for the incorporation of dissolved barium and molybdenum into the bivalve shell: an isotopic tracer approach in the juvenile Great Scallop (Pecten maximus). Marine Environmental Research 78: 15-25.
doi: 10.1016/j.marenvres.2012.03.006

13. Schöne B.R., Wanamaker Jr. A.D., Fiebig J., Thébault J. & Kreutz K. (2011). Annually resolved δ13Cshell chronologies of long-lived bivalve mollusks (Arctica islandica) reveal oceanic carbon dynamics in the temperate North Atlantic during recent centuries. Palaeogeography, Palaeoclimatology, Palaeoecology 302: 31-42.
doi: 10.1016/j.palaeo.2010.02.002

12. Schöne B., Zhang Z., Radermacher P., Thébault J., Jacob D., Nunn E. & Maurer A.F. (2011). Sr/Ca and Mg/Ca ratios of ontogenetically old, long-lived bivalve shells (Arctica islandica) and their function as paleotemperature proxies. Palaeogeography, Palaeoclimatology, Palaeoecology 302: 52-64.
doi: 10.1016/j.palaeo.2010.03.016

11. Chauvaud L., Thébault J., Clavier J., Lorrain A. & Strand Ø. (2011). What’s hiding behind ontogenetic δ13C variations in mollusk shells? New insights from the great scallop (Pecten maximus). Estuaries & Coasts 34: 211-220.
doi: 10.1007/s12237-010-9267-4

10. Barats A., Amouroux D., Pécheyran C., Chauvaud L., Thébault J. & Donard O.F.X. (2010). Spring molybdenum enrichment in scallop shells: a potential tracer of diatom productivity in temperate coastal environments (Brittany, NW France). Biogeosciences 7: 233-245.
doi: 10.5194/bg-7-233-2010

9. Thébault J., Chauvaud L., L’Helguen S., Clavier J., Barats A., Jacquet S., Pécheyran C. & Amouroux D. (2009). Barium and molybdenum records in bivalve shells: Geochemical proxies for phytoplankton dynamics in coastal environments? Limnology and Oceanography 54: 1002-1014.
doi: 10.4319/lo.2009.54.3.1002

8. Thébault J., Schöne B.R., Hallmann N., Barth M. & Nunn E.V. (2009). Investigation of Li/Ca variations in aragonitic shells of the ocean quahog Arctica islandica, northeast Iceland. Geochemistry, Geophysics, Geosystems 10: Q12008.
doi: 10.1029/2009GC002789

7. Barats A., Amouroux D., Chauvaud L., Pécheyran C., Lorrain A., Thébault J., Church T.M. & Donard O.F.X. (2009). High frequency barium profiles in shells of the Great Scallop Pecten maximus: a methodical long-term and multi-site survey in Western Europe. Biogeosciences 6: 157-170.
doi: 10.5194/bg-6-157-2009

6. Radermacher P., Schöne B.R., Gischler E., Oschmann W., Thébault J. & Fiebig J. (2009). Sclerochronology – a highly versatile tool for mariculture and reconstruction of life history traits of the queen conch, Strombus gigas (Gastropoda). Aquatic Living Resources 22: 307-318.
doi: 10.1051/alr/2009043

5. Thébault J., Schraga T.S., Cloern J.E. & Dunlavey E.G. (2008). Primary production and carrying capacity of former salt ponds after reconnection to San Francisco Bay. Wetlands 28: 841-851.
doi: 10.1672/07-190.1

4. Thébault J., Thouzeau G., Chauvaud L., Cantillánez M. & Avendaño M. (2008). Growth of Argopecten purpuratus (Mollusca: Bivalvia) on a natural bank in Northern Chile: sclerochronological record and environmental controls. Aquatic Living Resources 21: 45-55.
doi: 10.1051/alr:2008021

3. Thébault J., Chauvaud L., Clavier J., Guarini J., Dunbar R.B., Fichez R., Mucciarone D.A. & Morize E. (2007). Reconstruction of seasonal temperature variability in the tropical Pacific Ocean from the shell of the scallop, Comptopallium radula. Geochimica et Cosmochimica Acta 71: 918-928.
doi: 10.1016/j.gca.2006.10.017

2. Thébault J., Chauvaud L., Clavier J., Fichez R. & Morize E. (2006). Evidence of a 2-day periodicity of striae formation in the tropical scallop Comptopallium radula using calcein marking. Marine Biology 149: 257-267.
doi: 10.1007/s00227-005-0198-8

1. Fichez R., Adjeroud M., Bozec Y.-M., Breau L., Chancerelle Y., Chevillon C., Douillet P., Fernandez J.-M., Frouin P., Kulbicki M., Moreton B., Ouillon S., Payri C., Perez T., Sasal P. & Thébault J. (2005). A review of selected indicators of particle, nutrient and metal inputs in coral reef lagoon systems. Aquatic Living Resources 18: 125-147.
doi: 10.1051/alr:2005015

Participation in research projects