Biologist, ecologist, fisheries scientist



Lemar laboratory





My research activities at the Institute for Research for Development have focused since 1988 on tropical fish and aquatic ecosystems, in partnership with the scientific communities of South America (1988-1995) and Sub-Saharan Africa (since 1995). Between 2000 and 2010, I carried out scientific work on Marine Protected Areas in West Africa, in particular, in collaboration with stakeholders and populations. I have also studied biodiversity in the tropical Atlantic and participated in assessments conducted as part of the development of the IUCN Red List of East Atlantic fish species. I was a member of the IPBES panel of experts for the regional report “Africa” in which I was coordinator (CLA) of chapter 1. During my career, I have been principal investigator for several research projects, the last of which (ANR EPURE, 2012-2016) focused on heavy metal contamination in marine species in the northern Canary Current. Since July 2016, I am the head of the Laboratory of Environmental Marine Sciences (LEMAR), which is a joint reserch unit (UMR) of UBO, CNRS, IRD and Ifremer and I remain involved in several research projects on tropical marine ecology.

Afandi, I., Talba, S., Benhra, A., Benbrahim, S., Chfiri, R., Labonne, M., … Bouthir, F. Z. (2018). Trace metal distribution in pelagic fish species from the north-west African coast (Morocco). International Aquatic Research, 10(2), 191–205. Cite
Le Croizier, G., Lacroix, C., Artigaud, S., Le Floch, S., Raffray, J., Penicaud, V., … Tito-de-Morais, L. (2018). Significance of metallothioneins in differential cadmium accumulation kinetics between two marine fish species. Environmental Pollution, 236, 462–476. Cite
Simier, M., Ecoutin, J.-M., & Tito-de-Morais, L. (2019). The PPEAO experimental fishing dataset: Fish from West African estuaries, lagoons and reservoirs. Biodiversity Data Journal, 7(e31374), 54. Cite
Le Croizier, G., Schaal, G., Point, D., Le Loc’h, F., Machu, E., Fall, M., … Tito-de-Morais, L. (2019). Stable isotope analyses revealed the influence of foraging habitat on mercury accumulation in tropical coastal marine fish. Science of the Total Environment, 650, 2129–2140. Cite
Le Croizier, G., Lacroix, C., Artigaud, S., Le Floch, S., Munaron, J.-M., Raffray, J., … Tito-de-Morais, L. (2019). Metal subcellular partitioning determines excretion pathways and sensitivity to cadmium toxicity in two marine fish species. Chemosphere, 217, 754–762. Cite
Polidoro, B. A., Ralph, G. M., Strongin, K., Harvey, M., Carpenter, K. E., Arnold, R., … Williams, A. (2017). The status of marine biodiversity in the Eastern Central Atlantic (West and Central Africa). Aquatic Conservation: Marine and Freshwater Ecosystems. Cite
Sadio, O., Simier, M., Ecoutin, J.-M., Raffray, J., Laë, R., & Tito-de-Morais, L. (2015). Effect of a marine protected area on tropical estuarine fish assemblages: Comparison between protected and unprotected sites in Senegal. Ocean & Coastal Management, 116, 257–269. Cite
Kantoussan, J., Ecoutin, J. M., Laë, R., Thiaw, O. T., & Tito-de-Morais, L. (2018). A multimetric-index approach using fisheries data to assess fish assemblage structure in relation to salinity gradient in a tropical West African estuary. African Journal of Marine Science, 40(2), 175–185. Cite
Waeles, M., Planquette, H., Afandi, I., Delebecque, N., Bouthir, F., Donval, A., … Tito-de-Morais, L. (2016). Cadmium in the waters off South Morocco: Nature of particles hosting Cd and insights into the mechanisms fractionating Cd from phosphate. Journal of Geophysical Research: Oceans, 121(5), 3106–3120. Cite
Le Croizier, G., Schaal, G., Gallon, R., Fall, M., Le Grand, F., Munaron, J.-M., … Tito-de-Morais, L. (2016). Trophic ecology influence on metal bioaccumulation in marine fish: Inference from stable isotope and fatty acid analyses. Science of the Total Environment, 573, 83–95. Cite
Kaimuddin, A. H., Laë, R., & Tito-de-Morais, L. (2016). Fish Species in a Changing World: The Route and Timing of Species Migration between Tropical and Temperate Ecosystems in Eastern Atlantic. Frontiers in Marine Science. Marine Fisheries, Aquaculture and Living Resources, 162. Cite
Ecoutin, J.-M., Simier, M., Albaret, J.-J., Laë, R., Raffray, J., Sadio, O., & Tito-de-Morais, L. (2014). Ecological field experiment of short-term effects of fishing ban on fish assemblages in a tropical estuarine MPA. Ocean & Coastal Management, 100, 74–85. Retrieved from Cite
Colléter, M., Gascuel, D., Albouy, C., Francour, P., Tito-de-Morais, L., Valls, A., & Le Loc’h, F. (2014). Fishing inside or outside? A case studies analysis of potential spillover effect from marine protected areas, using food web models. Journal of Marine Systems, 139, 383–395. Retrieved from Cite
Ndour, I., Le Loc’h, F., Kantoussan, J., Thiaw, M., Diadhiou, H. D., Ecoutin, J.-M., … Thiaw, O. T. (2014). Changes in the trophic structure, abundance and species diversity of exploited fish assemblages in the artisanal fisheries of the northern coast, Senegal, West Africa. African Journal of Marine Science, 36(3), 361–368. Retrieved from Cite
Brochier, T., Ecoutin, J. M., Tito-de-Morais, L., Kaplan, D. M., & Laë, R. (2013). A multi-agent ecosystem model for studying changes in a tropical estuarine fish assemblage within a marine protected area. Aquatic Living Resources, 26(02), 147–158. Retrieved from Cite
Bodin, N., N’Gom-Kâ, R., Kâ, S., Thiaw, O. T., Tito-de-Morais, L., Le Loc’h, F., … Chiffoleau, J.-F. (2013). Assessment of trace metal contamination in mangrove ecosystems from Senegal, West Africa. Chemosphere, 90(2), 150–157. Retrieved from Cite
Kantoussan, J., Ecoutin, J. M., Simier, M., Tito-de-Morais, L., & Laë, R. (2012). Effects of salinity on fish assemblage structure: An evaluation based on taxonomic and functional approaches in the Casamance estuary (Senegal, West Africa). Estuarine, Coastal and Shelf Science, 113, 152–162. Retrieved from Cite
Guillard, J., Simier, M., Albaret, J.-J., Raffray, J., Sow, I., & Tito-de-Morais, L. (2012). Fish biomass estimates along estuaries: A comparison of vertical acoustic sampling at fixed stations and purse seine catches. Estuarine Coastal and Shelf Science, 107, 105–111. Cite
Azzoug, M., Carre, M., Chase, B. M., Deme, A., Lazar, A., Lazareth, C. E., … Tito-de-Morais, L. (2012). Positive precipitation-evaporation budget from AD 460 to 1090 in the Saloum Delta (Senegal) indicated by mollusk oxygen isotopes. Global and Planetary Change, 9899, 54–62. Cite
Colléter, M., Gascuel, D., Ecoutin, J.-M., & Tito-de-Morais, L. (2012). Modelling trophic flows in ecosystems to assess the efficiency of marine protected area (MPA), a case study on the coast of Senegal. Ecological Modelling, 232, 1–13. Cite
Faye, D., Tito-de-Morais, L., Raffray, J., Sadio, O., Thiaw, O. T., & Le Loc’h, F. (2011). Structure and seasonal variability of fish food webs in an estuarine tropical marine protected area (Senegal): Evidence from stable isotope analysis. Estuarine, Coastal and Shelf Science, 92(4), 607–617. Retrieved from Cite
Bodin, N., Ka, R., Le Loc’h, F., Raffray, J., Budzinski, H., Peluhet, L., & Tito-de-Morais, L. (2011). Are exploited mangrove molluscs exposed to persistent organic pollutant contamination in Senegal, West Africa? Chemosphere, 84(3), 318–327. Retrieved from Cite


Participation in research projects